Coreoidea Species File (Version 5.0/5.0)
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Taxa hierarchy
subfamily
Coreinae
Leach, 1815
tribe
Gonocerini
genus
Cletus
Stål, 1860
species
Cletus
pugnator
(Fabricius, 1787)
urn:lsid:Coreoidea.speciesfile.org:TaxonName:451963
Images: (8)
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specimen
specimen
specimen
Distribution:
Ecology:
Terrestrial.
Specimen records
are available.
Citations (30):
Fabricius. 1787. Mantissa insectorum sistens species nuper detectas adjectis synonymis, observationibus, descriptionibus, emendationibus 2:287
>> Note: Original description; STS [2 exx]: [India, Tamil Nadu], Tranquebar; ZMUC Copenhagen >>
Cimex
pugnator
urn:lsid:Coreoidea.speciesfile.org:TaxonName:456107
Gmelin. 1790. Caroli a Linné Systema Naturae 1(4):32137
>> Note: Description; loc. India (type loc.) >>
Cimex
pugnator
Latham. 1791. In Pennant. Indian Zoology. Second edition 130 >> Note: Listed; India >>
Cimex
pugnator
Fabricius. 1794. Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus 4:130
>> Note: Description; loc. India (type loc.) >>
Coreus
pugnator
urn:lsid:Coreoidea.speciesfile.org:TaxonName:456108
Weber. 1795. Nomenclator entomologicus secundum Entomologiam Systematicum illustr. Fabricii, adjectis speciebus recens detectis et varietatibus 152 >> Note: Listed >>
Coreus
pugnator
Turton. 1802. A general system of Nature, through the three grand kingdoms of animals, vegetables, and minerals, systematically divided into their several classes, orders, genera, species, and varieties, with their habitations, manners, economy, structure, and peculiarities, 1, Animal Kingdom; Vol. 2 Insects, Par 1802 632 >> Note: Description; loc. India (type loc.) >>
Cimex
pugnator
Fabricius. 1803. Systema Rhyngotorum secundum ordines, genera, speciesadjectis synonymis, locis, observationibus, descriptionibus. 197
>> Note: Description; loc. India (type loc.) >>
Coreus
pugnator
Donovan. 1807. In Rees [Ed.]. The Cyclopaedia; or, universal dictionary of arts, sciences, and literature 8 8 unnumbered pp >> Note: Description; loc. India (type loc.) >>
Coreus
pugnator
Burmeister. 1835. Handbuch der Entomologie 321
>> Note: Description; synonymy; Bengal >>
Hypselonotus
trigonus
Herrich-Schäffer. 1853. Die wanzenartigen Insecten 9(8-9):60, 107
>> Note: Catalogued; synonymy; India >>
Hypselonotus
trigonus
Dohrn. 1859. Catalogus Hemipterorum 28
>> Note: Catalogued; Bengal >>
Hypselonotus
trigonus
Signoret. 1861. In Montrouzier. Essai sur la faune entomologique de la Nouvelle-Calédonie (Balade) et des îles des Pins, Art, Lifu, etc. Hémiptères . Annales de la Société Entomologique de France 4 1:66 (footnote)
>> Note: As example of new subgenus Agonotomus attributed to Spinola >>
Gonocerus
(
Agonotomus
)
pugnator
urn:lsid:Coreoidea.speciesfile.org:TaxonName:456109
Stål. 1868. K. Svens. Vet.-Akad. Hand. 7(11):60
>> Note: Synonymy; description; compared with C. trigonus >>
Cletus
pugnator
urn:lsid:Coreoidea.speciesfile.org:TaxonName:451963
Stål. 1869. K. Svens. Vet.-Akad. Hand. (N.F.) 8(1):121 >> Note: Generic placement >>
Cletus
pugnator
Lethierry & Severin. 1894. Catalogue général des Hémiptères Tome II:67
>> Note: Catalogued (=pugnator, acutus, bistillatus); India, Sri Lanka, Philippines [Philippines record probably referable to trigonus] >>
Cletus
trigonus
Breddin. 1900. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 25(1):158
>> Note: Listed; rec. Borneo, recs Nias, Sumatra, Java >>
Cletus
pugnator
Distant. 1902. The fauna of British India, including Ceylon and Burma. Rhynchota 1 (Heteroptera) 394
>> Note: Synonymy (=pugnator, acutus, bistillatus); description; loc. India, recs Ceylon, North Borneo, Philippines [Philippines record probably referable to trigonus] >>
Cletus
trigonus
Breddin. 1905. Mitt. Hamb. Zool. xxii:118, 119
>> Note: Synonymy (=acutus); descriptive notes; loc. Java; compared with new variety Cletus punctulatus var. javanus; Cletus trigonus typicus >>
Cletus
trigonus
Breddin. 1909. Ann. Soc. Entomol. Belg. 53:289, 291
>> Note: Synonymy discussed (=Coreus bipunctatus Westwood, 1842; bistillatus, ‘Cletus trigonus var. pugnator Stål’ [Stål does not employ this name]); compared with C. trigonus in key; distribution: Indian, locs India (Madras, Sikkim), rec. Ceylon: common everywhere; compared with new species Cletus hoplomachus >>
Cletus
pugnator
Bergroth. 1913. Mémoires de la Société Entomologique de Belgique 22:152
>> Note: Catalogued (=bipunctatus Westwood, bistillatus Dohrn, inconspicuus Walker, signatus Walker, trigonus sensu Lethierry & Severin, 1894 in part); tropical Asia >>
Cletus
pugnator
Bergroth. 1915. Bombay J.Nat.Hist.Soc. 24:173
>> Note: Synonymy (=bipunctatus Westwood, which is not syn. of trigonus); loc. India >>
Cletus
pugnator
Blöte. 1935. Zoölogische Mededeelingen 18:207
>> Note: Locs India including 1 ‘cotype’ of Coreus bipunctatus Westwood, 1842 from ‘Poonah’ in RMNH Leiden >>
Cletus
pugnator
Dispons. 1963. Bull.Soc.Sci.nat.Laos 6:18-19 >> Note: Note on distribution: Oriental; species reaching Karakorum and Hindu Kush >>
Cletus
pugnator
Ahmad, Shadab & Khan. 1977. Supplement of the Entomological Society of Karachi 2:21 >> Note: Catalogued; locs Pakistan, loc. Bangladesh; on Oryza sativa and grass. >>
Cletus
trigonus
Josifov & Kerzhner. 1978. Fragmenta Faunistica (Warsaw) 23(9):161-162 >> Note: Synonymy (=bipunctatus Westwood (not bipunctatus Herrich-Schaeffer), not synonym of trigonus); compared with C. trigonus; recs Ceylon, India, Burma, S. China (Kweichow [=Guizhou]). >>
Cletus
pugnator
Ahmad. 1979. Supplement of the Entomological Society of Karachi 4(1):38 >> Note: Loc. Pakistan >>
Cletus
trigonus
Dang. 2000. Fauna of Vietnam 7:176, 270, 277 >> Note: Listed; identification; locs Vietnam; fig. 61 >>
Cletus
pugnator
Linnavuori. 2004. Acta Universitatis Carolinae, Biologia 48:280 >> Note: Habitat; loc. Iran, rec. Afghanistan (det. Kerzhner) >>
Cletus
pugnator
Dolling. 2006. In Aukema & Rieger [Ed.]. Catalogue of Heteroptera of the Palaearctic Region 5:77 >> Note: Catalogued (=bistillatus); notes; Palaearctic >>
Cletus
pugnator
Ghahari, Moulet, Linnavuori & Ostovan. 2012. Zootaxa 3519:8-9 >> Note: Distribution (Iran); >>
Cletus
pugnator
Scrutiny:
Dolling, 2011, Josifov & Kerzhner, 1978: 162 noted that
Cimex pugnator
Fabricius was synonymized with
Cimex trigonus
(Thunberg, 1783) by Stål, 1873: 78 but they regarded the two species as distinct. The record from China rests on material from Guizhou examined by them; Zheng & Dong, 1995 did not see any material from China and recorded
trigonus
from Sri Lanka and eastern India as well as much of SE Asia. Records from tropical Asia for
trigonus
and
pugnator
are inextricably confused. The names
Gonocerus acutus
Dallas, 1852 and
Cletus bistillatus
Dohrn, 1860b, cited here in synonymy with
Cletus trigonus
(Thunberg, 1783), may refer to this species.
Type specimen information:
There are multiple primary types, see specimen data for details.
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